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By Peter W. Hochachka

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Nidamental glands may also be present; these secrete an additional protective covering around the eggs. Octopods are unusual in having two oviducts. Fertilization may occur inside or outside the mantle cavity. A penis is never present and the spermatophore is conveyed to the female by a specialized and often very elaborate arm, the hectocotylus. In the pelagic octopod Argonauta the hectocotylus is detachable and moves about freely within the female mantle cavity. The two most dorsal arms are here greatly expanded and secrete a papery bivalved shell (not homologous with the shell of other molluscs) in which the female incubates her eggs and into which she can retreat when disturbed.

A ciliated region of the cecal wall separates out indigestible particles. The cephalopod digestive gland consists of "liver" and "pancreas," the former secreting into the cecum, the latter into the gizzard. The whole cephalopod alimentary canal is specialized for the rapid processing of food. C. Circulatory and Excretory Systems Although molluscs are coelomate, the true coelom is typically restricted to the cavities of the gonads and pericardium, and the main body cavity consists of a series of blood-filled lacunae that bathe the internal organs.

In the nudibranchs, tactile senses are especially important, and the whole naked dorsal surface becomes highly sensitive. Bivalves lack a head, and the major sense organs therefore come to lie mainly around the mantle margin or at the tips of the siphons. Mantle tentacles are tactile and chemosensory. They are especially well developed in the free-swimming pectinids where they may be associated with quite complex eyes. Statocysts occur in the foot but these are reduced or absent in attached forms such as oysters.

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